Memories of Pavel Kocourek (1948-2022)

By Petr Dolejs & Alena Kovaríková

Last year, just before Christmas, PaedDr. Pavel Kocourek passed away. Let us share with you two memories; one by his long-life friend Alena and the other by his colleague and let's say "a student" Petr.

Alena: I met Pavel Kocourek in the town of Týnec nad Sázavou for the first time. As a student at the secondary school, I was doing floristic research of the Cízov hill within the Natura semper viva competition. I needed scientific literature that was not available that time. I heard about a new teacher of biology at the primary school in Týnec nad Sázavou, so I asked him for help. Pavel lent me the books readily and he even invited me for an afternoon trip with his group of young naturalists. It was amazing how he behaved friendly to the children, how he was very dedicated to them and at the same time led them to independence. The children were showing him plants and animals they saw, and they were casting animal footprints. We all treated Pavel as an older, respected friend. We went to Panská skála Rock, observing nature, speaking about our hobbies and families. Pavel was proud of his three-year-old son, later I also met his wife Zdena and his parents. He visited me soon unexpectedly at my home, to look at my herbaria from the locality investigated. Pavel had a sense of humour. He noticed that I was living in a house with number 48; he told me that he comes from Prag where he also lived in a house of the same number and that he was born in 1948 and it is his lucky number. He offered me to cooperate as an instructor with his group of young naturalists, I attended two of his summer camps.
I had very similar interests and life opinions with Pavel. He was a soft man, who perceived atmosphere sensitively, had sense of the magic of the moment. He was discovering the beauty around him, he could catch it and forward his impression further. We were dealing with nature, often playing the guitar and singing, talking about history, culture, sport. Together with my husband Václav, we have been belonging to Pavel's closest friends for ages. During the last years, we were also taking part on the millipede study, Pavel taught us a lot and invited me to be a co-author of the Atlas of the Millipeds of the Czech Republic. We were performing together several-day research trips, my husband, as a reliable driver, often prepared the itineraries. Our last trip went to south-western Bohemia in October 2022. Noone could ever expect that this trip was our last one.

Petr: I knew Pavel in person since 2014, when I asked him for help with identification of millipedes collected in the Certovy schody Limestone quarry. We first met in my office in the National Museum of the Czech Republic, and I was surprised right from the beginning by his "ordinary" appearance, good mood and very friendly behaviour. Pavel identified the millipedes for me and because I was curious about the characters important for identification and wanted to learn more about the incredible millepedes, he started teaching me and became my informal mentor. I realized later that Pavel had caught the opportunity to have a new colleague in me. He liked being in touch with other people, other naturalists. We became friends soon and established a collaboration for further almost nine years during which we published 12 works and prepared four posters together.
I and Pavel complemented each other very well in our work. We were meeting either in my office but more often at Pavel's house in a small village Chynava in central Bohemia. I brought only my laptop with me, Pavel had served the rest (including lunch!). He was always very well prepared for our meetings (that was contrasting with Pavel's usual little cute absentmindedness); he always wrote down the questions he wished to discuss with me and was ready with all necessary documents. Pavel had admitted that it was easier for him painting than writing papers. I cannot paint and don't have so many experience with millipedes as Pavel had. Therefore, Pavel was giving me his ideas and I was translating and putting them on paper. We almost never argued. When we came to some critical point, we always explained to each other our opinions, and chose the best solution afterwards. I will never forget how we were preparing posters together. I knew what shall appear there from the scientific point of view, but I often had difficulties in how to put everything together graphically. And this was Pavel's role: he was choosing fonts, colours and correcting positions. I learnt from him how to present the science understandable, how to make it popular and easily accessible to public. Unfortunately, Pavel could not finish several projects he was working on. Together with Alena, we finished Pavel's key work - the Atlas of the Millipeds of the Czech Republic. However, three more manuscripts are still waiting for being completed. It will be my mission to finish what Pavel could not make.

Pavel Kocourek was born on 23 rd April 1948 in Prague. He was much influenced by his father Václav who taught him painting and knowing plants (Fig. 1). Pavel was always interested in nature, so it is not surprising that he initiated his studies at the secondary school specialized on chemistry and biology. He continued at the Faculty of Education of the Charles University in Prague (Fig. 2). His supervisor was a Czech diplopodologist Prof. Jaroslav Lang. Pavel defended his diploma thesis about the millipedes of Central Bohemia Region in 1970 (Fig. 3), got married with his first wife, Zdena, in 1971 (Fig. 4) and had three children, Pavel, Jana and Iva (Fig. 5). Then he was married (and divorced) two times more. In 1973, he published his first paper about millipedes of Lower Posázaví area and. .and left millipedes for almost three decades. Pavel Kocourek started his pedagogical career in 1970 and was teaching biology, chemistry, ecological practices and art at several basic schools (Fig. 6), and ecology and zoology at the Charles University (1998-2006). He was enjoying the work with children and young students. He was able not only to name everything in nature (beside animals and plants, he knew perfectly also lichens, minerals and fossils; KOCOUREK 1975, 1977), but also the stress the relationships between them (Fig. 7). This was something extraordinary that time given that ecology was not much "popular" by the communistic regime in the 1970's and 1980's. Pavel was also leading groups of young naturalists at the schools (that often caused him troubles for political reasons) and organized summer camps (Fig. 8), both in former Czechoslovakia and abroad, e.g. Germany (KOCOUREK & ZELENÝ 1980). In the 1990's (after the change of political regime), Pavel realized the lack of ecological knowledge, and established thus a system of environmental education at the basic schools (KOCOUREK 1991, 1992, 1993, 1996a, 1996b; PLACHEJDOVÁ & KOCOUREK 1996). The school, where he was a director that time, was the first schools that received a title "The Green school" (KOCOUREK 1998). He attended and organized several meetings for teachers of environmental education, established a network of ecological schools (KOCOUREK 1995, 1996c) and participated even on several textbook, either as an illustrator (STRIHAVKOVÁ & KUZMÍN 1971), coauthor (ANDRESKA et al. 1999; Fig. 9) or a lecturer (SLÉGL et al. 2002).

Pavel was also very active in nature protection. He proposed several natural reserves or natural monuments in the territory of now-a-day Czech Republic and created several natural educational trails (KOCOUREK 1990; FRIEDLOVÁ et al. 1991; SEDIVÁ et al. 1994; Fig. 10). He engaged in ecological competitions for students both as a co-organiser and leader of student teams. The teams led by him were very successful at those competitions and often ranked in the top positions. More information about Pavel's other activities can be found in previously published memories (DOLEJS et al. 2018; KOVARÍKOVÁ 2023; KOVARÍKOVÁ & DOLEJS 2023a, 2023b; DOLEJS & KOVARÍKOVÁ 2023a; SPRYNAR 2023).

Pavel returned to millipedes in 1998. After he oriented in changes that had occurred during the long period, he gathered information about species newly recorded in the Czech Republic (KOCOUREK 2001b, 2003) and analyzed the Czech millipede fauna (KOCOUREK 2001c). Then he initiated inventory research, namely in Prague (KOCOUREK 2001a, 2004), various protected areas (KOCOUREK 2007, 2013a, 2014; KOCOUREK & TAJOVSKÝ 2011, 2018; TAJOVSKÝ et al. 2022) and other places of interest (KOCOUREK et al. 2002; KOCOUREK 2018; KOCOUREK & DOLEJS 2018; DOLEJS & KOCOUREK 2019b). His long-term collecting efforts combined with a survey of historical resources resulted in a monograph about millipeds of Prague (KOCOUREK 2013b). This, as well as other Pavel's papers, were accompanied with his original drawings of alive millipedes and illustrations of their gonopods and taxonomically important characters. Pavel was an excellent painter (skilled after his father). He did not paint only millipedes but also landscape (Fig. 11) or experimental paintings and his painting were exposed at many exhibitions (Fig. 12), on only in the Czech Republic (e.g. in the Senate of the Czech Republic), but also abroad (Slovakia, Netherland, France, India, Japan). Many of us remember Pavel's hand-made greeting cards that he was sending for the New Year (Fig. 13). His outstanding drawings of millipedes accompanied most of his publications. He always paid attention to details; he was drawing the gonopods or telopods based on permanent slides that he had made. He always drew what he saw on the animal, often noting the natural variability. The exhibition of his paintings during the 16th ICM in Olomouc (2014) made deep impressions to all attendees (Fig. 14). The congresses belonged to Pavel's favourite entertainments. He attended the congresses in Poland (1999), South Africa (2002), Germany (2004) and the Czech Republic (2014; Fig. 15). He also planned to attend the congress in Hungary (2019) but finally could not come due to problems with his heart. He remembered well a lot of colleagues and their work, many of them became his friends.

Pavel participated also on Red Lists of Czech millipedes (KOCOUREK 2005; KOCOUREK & TAJOVSKÝ 2017a) and a list of alien species of Czech fauna (KOCOUREK 2006). During his research activities, he found 11 species new to the Czech Republic (KOCOUREK et al. 2017b; D OLEJS et al. 2019) - see also the list below. Pavel was able to do both science within the international teams (M OCK et al. 2016; NEFEDIEV et al. 2016) and presenting the millipedes to public (KOCOUREK 2017, 2021; KOCOUREK & DOLEJS 2019). On of the important part of Pavel's work was a collaboration with the National Museum of the Czech Republic. Here, he identified the millipedes from collection of several Czech naturalists (KOCOUREK & DOLEJS 2016; DOLEJS & KOCOUREK 2018, 2019a, 2020). The work in the museum resulted in deep cooperation with the first author of this article and led to two of the most important Pavel's works. In 2017, Pavel and his colleagues published a book about Czech millipedes (KOCOUREK et al. 2017a; CHOBOT 2017; T UF 2017; READ 2018). After getting familiar with yet unexplored parts of our country, Pavel initiated systematic research in those areas. We organized together many shorter or longer research trips or expeditions during the following five years (Fig. 16). Our data (and material that is all deposited in the National Museum of the Czech Republic) are base for an up-to-date Atlas of Distribution of Millipedes of the Czech Republic, published in 2023 (KOCOUREK et al. 2023). Unfortunately, Pavel will never see the result. We lost not only a colleague, "diplopodological guru", but also a very nice man, a friend (Fig. 17). We will continue in his work and work with the millipedes like Pavel will be together with us.

Millipede species discovered by Pavel as new for the Czech Republic:
Melogona broelemanni (Verhoeff, 1897): 17.x.1970, Veletov village, leg. P. Kocourek
Polydesmus angustus Latzel, 1884: 10.v.1971, Kolín town, leg. P. Kocourek
Haplogona oculodistincta (Verhoeff, 1893): 1.x.1999, Beskydy Mts., leg. P. Kocourek
Allajulus nitidus (Verhoeff, 1891): 13.x.-18.xii.2000, Prague, leg. P. Kocourek [L ANG 's (1952) record from the Hostice village is questionable according to G ULICKA (1985)] Cylindroiulus truncorum (Silvestri, 1896): 13.x.-18.xii.2000, Prague, leg. P. Kocourek
Cylindroiulus vulnerarius (Berlese, 1888): 13.x.-18.xii.2000, Prague, leg. P. Kocourek
Amphitomeus attemsi (Schubart, 1934): 2001, Prague, leg. P. Kocourek
Hungarosoma bokori Verhoeff, 1928: 13.x.2005, Moravian Karst PLA, leg. P. Kocourek & I. Skoumalová
Melogona transsylvanica (Verhoeff, 1897): 22.xi.2009, Krivoklátako PLA, leg. P. Kocourek
Haploglomeris multistriata (C. L. Koch, 1844): 24.vi.2021, Pasecná village, leg. P. Kocourek
Brachyiulus pusillus (Leach, 1816): 10.v.2021, Nezdice town, leg. P. Kocourek

Beside the species collected by Pavel himself, he also (co)published six new records of species collected by somebody else (KOCOUREK 2001b; DOLEJS & KOCOUREK 2018; KOCOUREK & DOLEJS 2019; DOLEJS et al. 2019):
- Brachyiulus lusitanus Verhoeff, 1898: 10.iv.1935, Brno, leg. A. Hoffer
- Rossiulus vilnensis (Jawlowski, 1925): 26.iv.-14.vi.1990, Zelezné hory Mts, leg. J. Buchar
- Propolydesmus germanicus (Verhoeff, 1896): 1996, Oblík NNR, leg. P. Moravec
- Cylindroiulus parisiorum (Brölemann & Verhoeff, 1896): 28.v.1998, Prague, leg. F. Stáhlavský
- Melogona gallica(Latzel, 1884): 28.v.-6.vii.2000, Lípový vrch Hill, leg. P. Vonicka
- Cylindrodesmus hirsutus Pocock, 1889: 20.viii.-7.x.2017, Prague, leg. V. Pesan

Diplopodological articles, chapters and monographs

DOLEJS, P.; KOCOUREK, P. (2018): Catalogue of the millipedes (Diplopoda) in the Hoffer's collection at the National Museum in Prague, Czech Republic. - Schubartiana 7: 1-6.
http://www.schubartiana.de/issues/pdf/vol7/Dolejs-2018- Catalogue_of_the_millipedes_in_the_Hoffer's_collection_Prague.pdf
DOLEJS, P.; KOCOUREK, P. (2019a): Bohumil Nemec and his millipede collection at the National Museum in Prague (Czechia), with notes on Craspedosoma rawlinsii simplex Nemec, 1896. - Schubartiana 8: 25-35. https://www.schubartiana.de/issues/pdf/vol8/Dolejs-2019- Nemec_millipede_collection_prague.pdf
DOLEJS, P.; KOCOUREK, P. (2019b): Príspevek k faune mnohonozek (Diplopoda) vybraných lokalit strední Moravy. Contribution to millipede fauna (Diplopoda) of selected localities of Central Moravia. - Zprávy Vlastivedného muzea v Olomouci 317: 98-107 (in Czech, with English abstract).
DOLEJS, P.; KOCOUREK, P. (2020): Catalogue of the millipedes (Diplopoda) in Miller's collection (Department of Zoology, National Museum, Prague, Czechia), part 2. - Journal of the National Museum (Prague), Natural History Series 189: 11-20. doi: http://dx.doi.org/ 10.37520/jnmpnhs.2020.003
DOLEJS, P.; KOCOUREK, P.; HULA, V. (2019): Cylindrodesmus hirsutus Pocock, 1889 new for Czechia (Diplopoda: Polydesmida: Haplodesmidae). - Schubartiana 8: 37-42. https://www.schubartiana.de/issues/pdf/vol8/Dolejs-2019- Cylindrodesmus_hirsutus_new_for_Czechia.pdf
KOCOUREK, P. (1970): Mnohonozky Stredoceského kraje [Millipedes of Central Bohemia Region]. - Thesis, Charles University, Czechoslovakia: 55 pp. (in Czech)
KOCOUREK, P. (1973): Mnohonozky Dolního Posázaví [Millipedes of Lower Posázaví]. - Sborník vlastivedných prací z Podblanicka 14: 78-93 (in Czech, with English summary).
KOCOUREK, P. (2001a): Pozoruhodné spolecenstvo mnohonozek Botanické zahrady UK (A remarkable community of millipedes of the Botanical Garden, Charles University). - Ziva 49 (3): 125-126 (in Czech, with English summary).
KOCOUREK, P. (2001b): Several new species of millipedes (Diplopoda) from the Czech Republic. - Acta Societatis Zoologicae Bohemicae 65 (2): 81-96. http://www.zoospol.cz/ixadmin/app/webroot/uploads/15-12-2017/2001/2001%202.pdf
KOCOUREK, P. (2001c): Druhová rozmanitost ceských mnohonozek v historickém kontextu. Species diversity of the Czech millipedes in the historical context. - Myriapodologica Czecho- Slovaca 1: 59-60 (in Czech, with English abstract).
KOCOUREK, P. (2003): New species of millipedes occurring in the Czech Republic: species discovered in the period 1970-2002. - African Invertebrates 44 (1): 199-202.
KOCOUREK, P. (2004): Mnohonozky - skrytá fauna v ZOO Praha (Millipedes - hidden fauna at the Prague Zoo). - Ziva 52 (4): 169-171 (in Czech, English summary).
http://ziva.avcr.cz/files/ziva/pdf/mnohonozky-skryta-fauna-v-zoo-praha.pdf KOCOUREK, P. (2005): Diplopoda (mnohonozky). - In: FARKAC, J.; KRÁL, D.; SKORPÍK, M. (eds): Cervený seznam ohrozených druhu Ceské republiky. Bezobratlí. Red list of threatened species in the Czech Republic. Invertebrates. Agentura ochrany prírody a krajiny CR, Praha: 760 pp. (in Czech and English).
KOCOUREK, P. (2006): Diplopoda - mnohonozky. - In: MLÍKOVSKÝ, J.; STÝBLO, P. (eds): Nepuvodní druhy fauny a flóry Ceské republiky [Alien species of fauna and flora of the Czech Republic]. Ceský svaz ochráncu prírody, Praha: 496 pp. (in Czech)
KOCOUREK, P. (2007): Mnohonozky (Diplopoda) chránené krajinné oblasti Ceský kras. Millipedes (Diplopoda) of the Protected Landscape Area Bohemian Karst. - Fragmenta Ioannea Collecta, Zoologica 7: 5-42 (in Czech, with English abstract).
KOCOUREK, P. (2013a): Mnohonozky [Millipedes]. - In: KARPAS R.; VISNÁK R.; VONICKA P. (eds): Jizerské hory, o raselinistích, kvetene a zvírene [Jizerské hory Mts, about peat bogs, plants and animals]. Nakladatelství RK, Liberec: 440 pp. (in Czech)
KOCOUREK, P. (2013b): Mnohonozky (Myriapoda: Diplopoda) Prahy. Millipedes (Myriapoda: Diplopoda) of Prague (Central Bohemia). - Natura Pragensis 21: 3-146 (in Czech, with English abstract and summary).
KOCOUREK, P. (2014): Mnohonozky (Myriapoda: Diplopoda) Jizerských hor, Frýdlantské pahorkatiny, Jestedského hrbetu a Liberecké kotliny (severní Cechy). Millipedes (Myriapoda: Diplopoda) of the Jizerské hory Mts, Frýdlantská pahorkatina Hills, Jestedský hrbet Ridge and Liberecká kotlina Basin (northern Bohemia). - Sborník Severoceského Muzea, Prírodní Vedy 32: 123-154 (in Czech, with English abstract and summary). https://www.muzeumlb.cz/IS/pu_data/send_files/Files/Sborniky/Sbornik_Prirodni_Vedy/ 32_2014/Kocourek_2014.pdf
KOCOUREK, P. (2017): Zajímaví pudní clenovci. Mnohonozky [Interesting soil arthropods. Millipedes]. - Krása naseho domova 59 (podzim/zima): 16-18 (in Czech).
KOCOUREK, P. (2018): Mnohonozky (Myriapoda: Diplopoda) Dolního Posázaví II. Millipedes (Myriapoda: Diplopoda) of the Lower Posázaví II. - Sborník vlastivedných prací z Podblanicka 52 (1-2): 7-44 (in Czech, with English abstract and summary).
KOCOUREK, P. (2021): Mnohonozky a príklady jejich sírení v Ceské republice [Millipedes and examples of their spreading in the Czech Republic]. - Podusták 1: 38-45 (in Czech).
KOCOUREK, P.; D OLEJS P. (2016): Catalogue of millipedes (Diplopoda) in Miller's collection (Department of Zoology, National Museum, Prague, Czechia). - Journal of the National Museum (Prague), Natural History Series 185 (4): 25-31. https://publikace.nm.cz/file/57848b49efab67a199d8aceac49684fe/19343/185-04- Kocourek-Millipedes.pdf
KOCOUREK, P.; DOLEJS, P. (2018): Mnohonozky (Diplopoda) Jezerského arboreta (okres Most, severozápadní Cechy). Millipedes (Diplopoda) of the Jezerí arboretum (Most district, Northwestern Bohemia). - Sborník Oblastního muzea v Moste, rada prírodovedná 39 ["2017"]: 98-105 (in Czech, English abstract). https://www.muzeummost.cz/data/stranky/Muzeum%20-%20Sbornik%20M17%20BLOK %20-%20Zoologie%201.pdf
KOCOUREK, P.; DOLEJS, P. (2019): Kolik druhu mnohonozek pribylo v Ceské republice v novém miléniu? (How many species of millipedes have arrived in the Czech Republic in the new millennium?). - Ziva 67 (4): 184-187 (in Czech, with English summary). https://ziva.avcr.cz/files/ziva/pdf/kolik-druhu-mnohonozek-pribylo-v-ceske-republice-v.pdf
KOCOUREK, P.; DOLEJS, P.; KOVARÍKOVÁ, A. (2023): Atlas rozsírení mnohonozek v Ceské republice. Atlas of the Millipedes of the Czech Republic. - Národní muzeum, Praha: 152 pp. (in Czech, with English summary). https://www.nm.cz/e-shop/atlas-rozsireni-mnohonozek-v-ceske- republice-atlas-of-the-millipedes-of-the-czech-republic
KOCOUREK, P.; FLASAROVÁ M.; FLASAR, I. (2000): Príspevky k faunistice mnohonozek (Diplopoda) severozápadních Cech. III. Beiträge zur Faunistik der Doppelfü?er (Diplopoda) Nordwestböhmens. III. - Sborník Okresního muzea v Moste, rada prírodovedná 22: 33-40 (in Czech, German summary).
KOCOUREK, P.; TAJOVSKÝ, K. (2011): Mnohonozky (Myriapoda, Diplopoda) CHKO a BR Krivoklátsko. Millipedes (Myriapoda: Diplopoda) of the Krivoklátsko PLA and BR. - Bohemia centralis 31: 285-300 (in Czech, English abstract). https://knihovna.nature.cz/media-viewer? rootDirectory=73402&origin=https%3A%2F%2Fknihovna.nature.cz%2Frecords%2Fd0d88889- aa01-4e76-a859-6b814c6e885b#!?file=30819
KOCOUREK, P.; TAJOVSKÝ, K. (2017a): Diplopoda (mnohonozky). - In: Hejda, R.; Farkac, J.; Chobot, K. (eds.): Cervený seznam ohrozených druhu Ceské republiky. Bezobratlí (Red List of Threatened Species of the Czech Republic. Invertebrates). - Príroda 36, Praha: 612 pp. (in Czech and English) https://www.priroda.nature.cz/index.php/priroda/article/view/30/59
KOCOUREK, P.; TAJOVSKÝ, K. (2018): Mnohonozky (Myriapoda: Diplopoda) CHKO Brdy. Millipedes (Myriapoda: Diplopoda) of the Brdy Protected Landscape Area. - Bohemia centralis 34: 187- 204 (in Czech, English abstract) https://knihovna.nature.cz/media-viewer?rootDirectory=73426&origin=https%3A%2F%2Fknihovna.nature.cz%2Frecords%2Fd133760f- df98-4b94-b4ae-ef0cc31bc2ee#!?file=30794
KOCOUREK, P.; TAJOVSKÝ, K.; DOLEJS, P. (2017a): Mnohonozky Ceské republiky - Prírucka pro urcování nasich druhu [Millipedes of the Czech Republic - Guide for identification of our species]. - Základní organizace Ceského svazu ochráncu prírody, Vlasim: 256 pp. (in Czech, with English abstract).
KOCOUREK, P.; TAJOVSKÝ, K.; DOLEJS, P. (2017b): New species of millipedes occurring in the Czech Republic: species discovered in the period 2003-2017. - Schubartiana 6: 27-30. http://www.schubartiana.de/issues/pdf/vol6/Kocourek-2017- New_species_Czech_Republic_2003-2017.pdf
M OCK, A.; TAJOVSKÝ, K.; ZUROVCOVÁ, M.; JAROSOVÁ, A.; KOCOUREK, P.; GRUBER, J.; ANGYAL, D.; SPELDA, J. (2016): Hungarosoma bokori Verhoeff, 1928 (Diplopoda: Chordeumatida): new insights into its taxonomy, systematics, molecular genetics, biogeography and ecology. - Zootaxa 4178 (2): 234-256. doi: http://doi.org/10.11646/zootaxa.4178.2.4
NEFEDIEV, P.S.; KOCOUREK, P.; NEFEDIEVA, J.S. (2016): The first record of Polydesmus inconstans Latzel, 1884 (Diplopoda: Polydesmida: Polydesmidae) in the Asian part of Russia. - Arthropoda Selecta 25 (1): 19-21. http://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_1_019_021_Nefediev_for_In et.pdf
TAJOVSKÝ, K.; M ACHAC, O.; KOCOUREK, P.; STASIOV, S.; DOLEJS, P.; ZDRÁHALOVÁ, H.; TUF, I.H. (2022): Výsledky myriapodologické a isopodologické exkurze v CHKO Zelezné hory. Results of myriapodological and isopodological excursion in the Zelezné hory Protected Landscape Area. - Východoceský sborník prírodovedný - práce a studie 28: 85-92 (in Czech, with English summary).
See CIM Newsletter 2023 (p. 78-94) for full obituary with figures